Harpalus

Harpalus es un género de coleópteros adéfagos de la familia Carabidae.^[1]​

Harpalus
Harpalus affinis
Taxonomía
Reino:	Animalia
Filo:	Arthropoda
Clase:	Insecta
Orden:	Coleoptera
Suborden:	Adephaga
Familia:	Carabidae
Subfamilia:	Harpalinae
Tribu:	Harpalini
Subtribu:	Harpalina
Género:	Harpalus Latreille, 1802
Especies
Ver texto
[editar datos en Wikidata]

Principalmente holártico con unas pocas especies en la región afrotropical. Hay 415 especies.^[2]​

Algunas especies

Comprende las siguientes especies:^[3]​

• Harpalus actiosus Casey, 1914
• Harpalus acupalpoides Reitter, 1900
• Harpalus adenticulatus Huang, 1992
• Harpalus aeneipennis Faldermann, 1836
• Harpalus aenigma (Tschitscherine, 1897)
• Harpalus aequicollis Motschulsky, 1844
• Harpalus aesculanus Pantel, 1888
• Harpalus affinis (Schrank, 1781)
• Harpalus agakhaniantzi Mikhailov, 1972
• Harpalus akinini Tschitscherine, 1895
• Harpalus alajensis Tschitscherine, 1898
• Harpalus albanicus Reitter, 1900
• Harpalus alexandrae Kataev, 1990
• Harpalus alexeevi Kataev, 1990
• Harpalus alienus Bates, 1878
• Harpalus alpivagus Tschitscherine, 1899
• Harpalus amarellus Bates, 1891
• Harpalus amariformis Motschulsky, 1844
• Harpalus amplicollis Menetries, 1848
• Harpalus amputatus Say, 1830
• Harpalus anatolicus Tschitscherine, 1898
• Harpalus angulatus Putzeys, 1878
• Harpalus angustipennis Boheman, 1848
• Harpalus angustitarsis Reitter, 1887
• Harpalus animosus Casey, 1924
• Harpalus anisodactyliformis Solsky, 1874
• Harpalus antonowi Tschitscherine, 1898
• Harpalus anxioides Kataev, 1991
• Harpalus anxius (Duftschmid, 1812)
• Harpalus aogashimensis (Habu, 1957)
• Harpalus apache Kataev, 2010
• Harpalus araraticus Mlynar, 1979
• Harpalus arcuatus Tschitscherine, 1898
• Harpalus arnoldii Kataev, 1988
• Harpalus ascetes Kataev & Wrase, 1997
• Harpalus asemus Basilewsky, 1946
• Harpalus asphaltinus Roth, 1851
• Harpalus aterrimus Casey, 1914
• Harpalus atratus Latreille, 1804
• Harpalus atrichatus Hatch, 1949
• Harpalus atripes Casey, 1914
• Harpalus attenuatus Stephens, 1828
• Harpalus australasiae Dejean, 1829
• Harpalus autumnalis Duftschmid, 1812
• Harpalus azumai Habu, 1968
• Harpalus babai Habu, 1973
• Harpalus badakschanus Jedlicka, 1956
• Harpalus baleensis Clarke, 1973
• Harpalus balli Noonan, 1991
• Harpalus basanicus J.R. Sahlberg, 1911
• Harpalus basilewskyi Facchini, 2003
• Harpalus basuto Basilewsky, 1958
• Harpalus bellieri Reiche, 1861
• Harpalus beneshi Kataev & Wrase, 1997
• Harpalus bitinctus Jeannel, 1948
• Harpalus brachypterus Tschitcherine, 1898
• Harpalus brevicornis Germar, 1824
• Harpalus brevis Motschiulsky, 1844
• Harpalus breviusculus Chaudoir, 1846
• Harpalus brunnipes Dejean, 1829
• Harpalus bucharicus Tschitscherine, 1898
• Harpalus bungii Chaudoir, 1844
• Harpalus caeruleatus Bates, 1878
• Harpalus caiphus Reiche & Saulcy, 1855
• Harpalus calathoides Motschulsky, 1844
• Harpalus calceatus (Duftschmid, 1812)
• Harpalus caliginosus (Fabricius, 1775)
• Harpalus capicola Dejean, 1829
• Harpalus capito A.Morawitz, 1862
• Harpalus cardoni Antoine, 1922
• Harpalus caspius (Steven, 1806)
• Harpalus cautus Dejean, 1829
• Harpalus cephalotes Fairmaire & Laboulbene, 1854
• Harpalus chalcentus Bates, 1873
• Harpalus chalcopterus Jeannel, 1948
• Harpalus chasanensis Lafer, 1989
• Harpalus chengjiangensis Huang, 1993
• Harpalus chiloschizontus Huang, 1995
• Harpalus chobautianus Lutshnik, 1922
• Harpalus chrysopus Reitter, 1887
• Harpalus cilihumerus Huang, Hu & Sun, 1994
• Harpalus circumpunctatus Chaudoir, 1846
• Harpalus cisteloides Motschulsky, 1844
• Harpalus cohni Ball, 1972
• Harpalus colchicus Motschulsky, 1850
• Harpalus compar Leconte, 1848
• Harpalus compressus Motschulsky, 1844
• Harpalus contemptus Dejean, 1829
• Harpalus convexus Faldermann, 1836
• Harpalus cordifer Notman, 1919
• Harpalus cordifer Notman, 1919
• Harpalus coreanus (Tschitscherine, 1895)
• Harpalus corporosus Motschulsky, 1861
• Harpalus corpulentus (Casey, 1914)
• Harpalus corrugatus Basilewsky, 1958
• Harpalus crates Bates, 1883
• Harpalus cupreus Dejean, 1829
• Harpalus cursorius Peringuey, 1896
• Harpalus cyanopterus Tschitscherine, 1897
• Harpalus cyclogonus Chaudoir, 1844
• Harpalus cyrtonotoides Notman, 1919
• Harpalus danieli Reitter, 1900
• Harpalus davidi (Tschitscherine, 1897)
• Harpalus davidianus Tschitscherine, 1903
• Harpalus decipiens Dejean, 1829
• Harpalus defector Peringuey, 1896
• Harpalus demelti Korge, 1962
• Harpalus desertus Leconte, 1859
• Harpalus diligens Tschitscherine, 1898
• Harpalus dimidiatus (P. Rossi, 1790)
• Harpalus disaogashimensis Huang In Huang & Zhang, 1995
• Harpalus disimuciulus Huang, Lei, Yan & Hu, 1996
• Harpalus dispar Dejean, 1829
• Harpalus dissitus Antoine, 1931
• Harpalus distinguendus (Duftschmid, 1812)
• Harpalus diversicollis Basilewsky, 1958
• Harpalus dubius Boheman, 1848
• Harpalus durangoensis Bates, 1891
• Harpalus ebeninus Heyden, 1870
• Harpalus eberlovi O. Berlov, 1996
• Harpalus egenus Dejean, 1829
• Harpalus egorovi Lafer, 1989
• Harpalus ellipsis Leconte, 1848
• Harpalus eous Tschitscherine, 1901
• Harpalus eraticus Say, 1823
• Harpalus erosus Mannerheim, 1825
• Harpalus erythropus Dejean, 1829
• Harpalus euchlorus Menetries, 1839
• Harpalus exiguus Boheman, 1848
• Harpalus famelicus Tschitscherine, 1898
• Harpalus farkaci Kataev & Wrase, 1995
• Harpalus faunus Say, 1823
• Harpalus ferghanensis Tschitscherine, 1899
• Harpalus fimetarius Dejean, 1829
• Harpalus flavescens Piller & Mitterpacher, 1783
• Harpalus flavicornis Dejean, 1829
• Harpalus fokienensis Schauberger, 1930
• Harpalus foveiger Tschitscherine, 1895
• Harpalus franzi Mateu, 1954
• Harpalus frater Chaudoir, 1876
• Harpalus fraternus Leconte, 1852
• Harpalus froelichii Sturm, 1818
• Harpalus fukiensis Jedlicka, 1956
• Harpalus fulgens Csiki, 1932
• Harpalus fulvicornis Thunberg, 1806
• Harpalus fulvilabris Mannerheim, 1853
• Harpalus fulvipennis Chaudoir, 1843
• Harpalus fulvus Dejean, 1829
• Harpalus fuscicornis Menetries, 1832
• Harpalus fuscipalpis Sturm, 1818
• Harpalus fuscipennis Wiedemann, 1825
• Harpalus fuscoaeneus Dejean, 1829
• Harpalus ganssuensis Semenov, 1889
• Harpalus giacomazzoi Kataev & Wrase, 1996
• Harpalus gilgil Basilewsky, 1946
• Harpalus gisellae Csiki, 1932
• Harpalus glasunovi Kataev, 1987
• Harpalus gravis Leconte, 1858
• Harpalus gregoryi Alluaud, 1917
• Harpalus grilli Kataev, 2001
• Harpalus griseus (Panzer, 1796)
• Harpalus hartmanni Kataev, 2001
• Harpalus hatchi Ball & Anderson, 1962
• Harpalus hauserianus Schauberger, 1929
• Harpalus herbivagus Say, 1823
• Harpalus heyrovskyi Jedlicka, 1928
• Harpalus hiekei Kataev & Wrase, 2010
• Harpalus hirtipes Panzer, 1796
• Harpalus honestus (Dufischmid, 1812)
• Harpalus hospes Sturm, 1818
• Harpalus huachuca Ball, 1972
• Harpalus hybridus Boheman, 1848
• Harpalus hypogeomysis Huang, 1993
• Harpalus idiotus Bates, 1889
• Harpalus imerinae Jeannel, 1948
• Harpalus impressicollis Jeannel, 1948
• Harpalus impressus Roth, 1851
• Harpalus inconcinnus Chaudoir, 1876
• Harpalus indianus Csiki, 1932
• Harpalus indicola Bates, 1878
• Harpalus indicus Bates, 1891
• Harpalus indigens Casey, 1924
• Harpalus inexspectatus Kataev, 1989
• Harpalus ingenuus Tschitscherine, 1898
• Harpalus innocuus Leconte, 1863
• Harpalus italus Schaum, 1860
• Harpalus janinae Jeanne, 1984
• Harpalus jeanneli Basilewsky, 1946
• Harpalus jordanus Jedlicka, 1964
• Harpalus jureceki (Jedlicka, 1928)
• Harpalus kabakianus Kataev, 1988
• Harpalus kabakovi Kataev, 1987
• Harpalus kadleci Kataev & Wrase, 1995
• Harpalus kadyrbekovi Kataev, 1988
• Harpalus kagyzmanicus Kataev, 1984
• Harpalus kailiensis Huang, 1992
• Harpalus kandaharensis Jedlicka, 1956
• Harpalus karamani Apfelbeck, 1902
• Harpalus karokorum Jedlicka, 1958
• Harpalus katiae F.Battoni, 1985
• Harpalus kazanensis Jedlicka, 1958
• Harpalus kaznakovi Kataev & Wrase, 1997
• Harpalus kibonoti Alluaud, 1926
• Harpalus kirgisicus Motschulsky, 1844
• Harpalus kiritshenkoi Kataev, 1990
• Harpalus klapperichi Jedlicka, 1956
• Harpalus kmecoi Facchini, 2003
• Harpalus koshiensis Kirschenhofer, 1992
• Harpalus kozlovi Kataev, 1993
• Harpalus kryzhanovskii Kataev, 1988
• Harpalus kunarensis Kataev, 1993
• Harpalus laetus Reiche, 1843
• Harpalus laevipes Zetterstedt, 1828
• Harpalus lama Kataev & Wrase, 1997
• Harpalus lampronotus Jeannel, 1948
• Harpalus lateralis Dejean, 1829
• Harpalus laticeps Leconte, 1850
• Harpalus latus (Linnaeus, 1758)
• Harpalus lederi Tschitscherne, 1899
• Harpalus lethierryi Reiche, 1860
• Harpalus lewisii Leconte, 1865
• Harpalus liobasis Chaudoir, 1868
• Harpalus litigiosus Dejean, 1829
• Harpalus longicollis Leconte, 1848
• Harpalus longihornus Lei & Huang, 1997
• Harpalus longipalmatus Mordkovitsh, 1969
• Harpalus lopezi Serrano & Lencina, 2008
• Harpalus lugubris Boheman, 1848
• Harpalus lumbaris Mannerheim, 1825
• Harpalus luteicornis Dufischmid, 1812
• Harpalus lutshniki Schauberger, 1932
• Harpalus macronotus Tschitscherine, 1893
• Harpalus madagascariensis Dejean, 1831
• Harpalus mairei Peyerimhoff, 1928
• Harpalus major Motschulsky, 1850
• Harpalus makhekensis Basilewsky, 1958
• Harpalus manas Kataev, 1990
• Harpalus marginellus Gyllenhal, 1827
• Harpalus mariae Kataev, 1997
• Harpalus martini Vandyke, 1926
• Harpalus masoreoides Bates, 1878
• Harpalus massarti Burgeon, 1935
• Harpalus mauritanicus Gaubil, 1844
• Harpalus medvedevi Kataev, 2006
• Harpalus megacephalus Leconte, 1848
• Harpalus meghalayensis Kataev, 2001
• Harpalus melancholicus Dejean, 1829
• Harpalus melaneus Bates, 1878
• Harpalus meridianus Andrewes, 1923
• Harpalus metallinus Menetries, 1836
• Harpalus metarsius Andrewes, 1930
• Harpalus meteorus Basilewsky, 1946
• Harpalus michaili Kataev, 1990
• Harpalus michailovi Kataev, 1987
• Harpalus microdemas Schauberger, 1932
• Harpalus microthorax Motschulsky, 1849
• Harpalus miles Peringuey, 1896
• Harpalus minutulus Kataev & Liang, 2004
• Harpalus mitridati Pliginskiy, 1915
• Harpalus mlynari Kataev, 1990
• Harpalus modestus Dejean, 1829
• Harpalus morvani Kataev, 2001
• Harpalus muciulus Huang, 1992
• Harpalus nanniscus Peringuey, 1896
• Harpalus natalensis Boheman, 1848
• Harpalus natalicus Peringuey, 1896
• Harpalus neglectus Audinet-Serville, 1821
• Harpalus nevadensis K. Daniel & J. Daniel, 1898
• Harpalus nigrans A. Morawitz, 1862
• Harpalus nigritarsis C.R. Sahlberg, 1827
• Harpalus numidicus Bedel, 1893
• Harpalus nyassicus Basilewsky, 1946
• Harpalus obliquus G.Horn, 1880
• Harpalus oblitus Dejean, 1829
• Harpalus obnixus Casey, 1924
• Harpalus obtusiusculus Jeannel, 1948
• Harpalus ochropus Kirby, 1837
• Harpalus oodioides Dejean, 1829
• Harpalus opacipennis (Haldemann, 1843)
• Harpalus optabilis Dejean, 1829
• Harpalus ovtshinnikovi Kataev, 1990
• Harpalus pachys Jeannel, 1948
• Harpalus paivanus Wollaston, 1867
• Harpalus pallidipennis A. Morawitz, 1862
• Harpalus pallipes Chaudoir, 1837
• Harpalus parasinuatus Kataev & Liang, 2007
• Harpalus paratus Casey, 1924
• Harpalus parvulus Dejean, 1829
• Harpalus pastor Motschulsky, 1844
• Harpalus penglainus Huang, Hu & Sun, 1994
• Harpalus pensylvanicus (Degeer, 1774)
• Harpalus periglabellus Huang, 1992
• Harpalus perlucens Bates, 1878
• Harpalus petri Tschitscherine, 1902
• Harpalus picipennis Duftschmid, 1812
• Harpalus pilosus Huang, 1995
• Harpalus plancyi Tschitscherine, 1897
• Harpalus platynotus Bates, 1873
• Harpalus plenalis Casey, 1914
• Harpalus politus Dejean, 1829
• Harpalus polyglyptus Schaum, 1862
• Harpalus poncei Will, 2001 Usa (Florida)
• Harpalus potanini Tschitscherine, 1906
• Harpalus poussereaui Pierre & Mauro, 2010
• Harpalus praecurrens Schauberger, 1934
• Harpalus praticola Bates, 1891
• Harpalus procognatus Lorenz, 1998
• Harpalus progrediens Schauberger, 1922
• Harpalus prosperus Basilewsky, 1972
• Harpalus protractus Casey, 1914
• Harpalus providens Casey, 1914
• Harpalus pseudohauserianus Kataev, 2001
• Harpalus pseudophonoides Schauberger, 1930
• Harpalus pseudoserripes Reitter, 1900
• Harpalus pseudotinctulus Schauberger, 1932
• Harpalus pterostichus Reitter, 1900
• Harpalus puetzi Kataev & Wrase, 1997
• Harpalus pulchirinulus Reitter, 1900
• Harpalus pulvinatus Menetries, 1848
• Harpalus pumilus Sturm, 1818
• Harpalus punctatostriatus Dejean, 1829
• Harpalus puncticeps (Casey, 1914)
• Harpalus punctipennis Mulsant, 1852
• Harpalus pusillus Motschulsky, 1850
• Harpalus pygmaeus Dejean, 1829
• Harpalus quadratus Chaudoir, 1846
• Harpalus raphaili Kataev, 1997
• Harpalus reflexus Putzeys, 1878
• Harpalus regressus (Casey, 1914)
• Harpalus remboides Solsky, 1874
• Harpalus retractus Leconte, 1863
• Harpalus reversus Casey, 1924
• Harpalus rivalsi Jeannel, 1948
• Harpalus roninus Bates, 1873
• Harpalus rotundus Facchini, 2003
• Harpalus rougemonti Clarke, 1973
• Harpalus rubefactus Bates, 1873
• Harpalus rubripes Dufischmid, 1812
• Harpalus rufipalpis Sturm, 1818
• Harpalus rufipes (Degeer, 1774)
• Harpalus rufiscapus Gebler, 1833
• Harpalus rufitarsoides Schauberger, 1934
• Harpalus rufomarginatus (Boheman, 1848)
• Harpalus rumelicus Apfelbeck, 1904
• Harpalus salinulus Reitter, 1900
• Harpalus salinus Dejean, 1829
• Harpalus sanctaehelenae Basilewsky, 1972
• Harpalus saxicola Dejean, 1829
• Harpalus scabripectus Huang, Hu & Sun, 1994
• Harpalus schaumii Wollaston, 1864
• Harpalus semenowi Tschitscherine, 1901
• Harpalus sericatus Tschitscherine, 1906
• Harpalus serripes Quensel, 1806
• Harpalus servus Duftschmid, 1812
• Harpalus setiporus Reitter, 1894
• Harpalus seyrigi Jeannel, 1948
• Harpalus siculus Dejean, 1829
• Harpalus signaticornis (Duftschmid, 1812)
• Harpalus silipes Dejean, 1831
• Harpalus simplicidens Schauberger, 1929
• Harpalus singularis Tschitscherine, 1906
• Harpalus sinicus Hope, 1845
• Harpalus sinuatipennis Jeannel, 1948
• Harpalus sinuatus Tschitscherine, 1893
• Harpalus smaragdinus Duftschmid, 1812
• Harpalus smyrnensis Heyden, 1888
• Harpalus solitaris Dejean, 1829
• Harpalus somereni Basilewsky, 1946
• Harpalus somnulentus Dejean, 1829
• Harpalus spadiceus Dejean, 1829
• Harpalus spretus Peringuey, 1896
• Harpalus spurius Peringuey, 1896
• Harpalus stephani Ball, 1972
• Harpalus stevenii Dejean, 1829
• Harpalus stoetznerianus Schauberger, 1932
• Harpalus strenuus Tschitscherine, 1898
• Harpalus stricticollis Jeannel, 1948
• Harpalus subaeneus Boheman, 1848
• Harpalus subcylindricus Dejean, 1829
• Harpalus subphaedrus Basilewsky, 1946
• Harpalus subtruncatus Chaudoir, 1846
• Harpalus suensoni Kataev, 1997
• Harpalus sulphuripes Germar, 1824
• Harpalus sundaicus Schauberger, 1933
• Harpalus sushenicus Kataev, 1990
• Harpalus suturangulus Reitter, 1887
• Harpalus szalliesi Kataev & Wrase, 1995
• Harpalus taciturnus Dejean, 1829
• Harpalus tadorcus Ball, 1972
• Harpalus tangutorum Kataev, 1993
• Harpalus tardus Panzer, 1796
• Harpalus tarsalis Mannenheim, 1825
• Harpalus tenebrosus Dejean, 1829
• Harpalus terrestris Motschulsky, 1844
• Harpalus texanus Casey, 1914
• Harpalus tianschanicus Semenov, 1889
• Harpalus tibeticus Andrewes, 1930
• Harpalus tichonis Jakobson, 1907
• Harpalus tinctulus Bates, 1873
• Harpalus tiridates Reitter, 1900
• Harpalus tithonus Reitter, 1900
• Harpalus torosensis Jedlicka, 1961
• Harpalus torridoides Reitter, 1900
• Harpalus trichophorus Tschitscherine, 1897
• Harpalus tridens A.Morawitz, 1862
• Harpalus triseriatus A. Fleischer, 1897
• Harpalus turcicus Jedlicka, 1958
• Harpalus turmalinus Erichson, 1847
• Harpalus udege Lafer, 1989
• Harpalus uniformis Motschulsky, 1844
• Harpalus ussuricus Mlynar, 1979
• Harpalus ussuriensis Chaudoir, 1863
• Harpalus vanemdeni Schauberger, 1932
• Harpalus venator Boheman, 1848
• Harpalus ventralis Leconte, 1848
• Harpalus vereschaginae Kataev, 1988
• Harpalus vernicosus Kataev & Liang, 2007
• Harpalus viridanus Motschulsky, 1844
• Harpalus viridicupreus Reiche, 1843
• Harpalus vittatus Gebler, 1833
• Harpalus wadiensis Jedlicka, 1964
• Harpalus wohlberedti Emden & Schauberger, 1932
• Harpalus xanthopus Gemminger & Harold, 1868
• Harpalus xinjiangensis Huang, Hu & Sun, 1994
• Harpalus yinchuanensis Huang, 1993
• Harpalus zabroides Dejean, 1829
• Harpalus zhdankoi Kataev, 1990

Referencias

1. Harpalus en BioLib
2. Bugguide.net. Genus Harpalus
3. . Carabidae of the World. 2011. Archivado desde el original el 15 de septiembre de 2012. Consultado el 1 de julio de 2011. 

Enlaces externos

• ION
• Nomenclator Zoologicus

•   Datos: Q1766785
•   Multimedia: Harpalus
•   Especies: Harpalus